CHAPTER FIVE
DISCUSSION
GROWTH
Th€ Srowrlr
perfomde of l/,
rvd siSnificdl iDlllcnced
by thc
nhiz, Cirhi"us tuigdlo
eo
and
Catld catla
ldels offadios S.eeppa.nd D€vd,j ( |991) tepoded
$ar grcwtn Etc wG 3iSnificmtly dep.nd.nt on fcdi.S levels TlE linal Ncidls
fish
A.ci6
0a
high.r sl 4 pqenr
l.v.l of f..dhs thd .t
Kh'n !! c!. (l t9l) cponed lhal mjor cs'p ll.s.rlins
percent
of body
willtt
of Renuta6dnyq and
Icvel equivslefl to 5
had
l'vcl ol lc'di;&
ol lhe pFsdl studv suppon thc r$ulls
lwice daily The
'indings
vdgher (1e86) lh.y rporLd
tnal
hirl nk offeding tbov'
Frenr ollh. innial bodv Eiglll of Mjd
ch@El col fish (Cotlin
Fr6l
8E!i6 wishl iah sbcn red 4
son {tEl sinilr ,indingr h.vc b@r GFdcd wilh 4a.di
4
feeding codd bc duc lo lhe
also suppon $e
2
41, 1986)- Thc Poor gowlh
1o
cary6
Mv
not be
lhe
uetu!
le€ls ol f"di'g for tlE
obsed
al 2
FGnt
levcl
of
fed slpply bclow iti .eql. Tte finding! oflh' pEsent slldv
Elulc orCho s! a! rla?6l Th.v shovtd lhal
reduction rn tlE lercl
f..diog Esnhsd in lovg fDal wichl ol njnbow lrcd Tn 6ulls su8ael lhat
level of feeding ie moE suihbl. fo. betlq fsh
Th? vlrittions in wights ot
diers i.e., prcGin lcv.ls wben
$ber€d tn€ vdiatio6
oflttr*
r.E
(he
4
of
Frccnt
go$r[.
lrsh 3pccr6
w d rsnilianl wi'h E8; b
6rio oftninMllo lcg.toble polein wa 70J0
(Phase
l)'
sladtlially signilicdt wh.n €tio o'eimdl rc veselabl'
Itt
pol.in wd
10:70
proeh ih fish
(Phe II). lllis sh.sed rhd r adaid
rc].
dier pldy a
in smwlh ra$
. Undcr phas I, th. hishs
rof
ik ad cano
.zna
ad
Ca|o catla
rfEt the
eflet ofdi.B
body
to Ege|ablc
lEietrs s.r. obsvcd in.usc ol Zdrd,
it rE hishs in c* oI
lsf/,) pml.ir di.r .r 2 FE6t lev.l ol f..dilg.
nuihu
w
{imal
olrhEc tsh sp@r.s
q be (40e4) prolcin di.t {heB
Cithinus ,tigola oh high
l€vel of f..ding, ihe
finil
ratio of
Ar 4
nnd body w.ighb of Zabc, rohita, Ctnhnns dtigal4
on nEdiuD, li8h @d loe prcLin dieb, r.sp.clively.
aor
p@dt
CirrihB krisala &d
Catlo .oda
fading d.e4. tor Lab.o rohita. Utrd.r Ph& ll, th.
w
T
his shors
simih d bod
Mimm
lin
l wisirls
levels
of
of
lzr@
rchita wd Cithlnus nrtgala w.rc on higl Folein diel and of Cat?a @r. on low protein
diet at 2 perenl lwcl of feding whe@s
sc
of 2
p.e
vrirtios
phase II
.t 4 p€rcql l.v€l of
feeding, rh.
tr.rd ms lhe
lc!.|of a..diIg.
in rh.
nighl b. dlc
t
{d
of fist body
to rh. chanee in
witt
ot
bb.o nkta 6dd llve | .nd
6lio of rrinal lo wgelabl€ pol.in.
In
phas II, this
fishspecisntghtlEveprefercd!oeathightrclcindielinln.abseneofhiShprcponion
of
sin.l
porein in thc dicl.
A5 r€gnd3
oqall greert perfor@ ,JMA tbc
d3h
!Fci.3 uDdd pn!* I
Lob.o rohita aod Cnrhh6 tut'gala hav. ihown beltq grc*th p.rfom&c.
.o,/a on loq and high protein diell. Whcrca lnder phde
rrisata \|6 berr.t rl\8
rhat Labeo
The
los
rcltih ad
I
th{
lhc B@$1h ol
Crrri"6
Labeo rohitq atd Ca a cd a @ bign Wnr.i. dict. This
Cir
ins urg4L hE€
fi.
ond hish dicro.y prctein lev.ls pdsibly
Cala
EEded
poLnlial lo gow .xcluiv.ly on di.l.
h. @middd fd
Labeo rohra
^ntl
ltt
Chrl'nn6 nDigala stEn cttrurr,l und.r
rh,r
th* lhF
6sh sFcic3 De.d.n
Th. pe*nr Eurrs sub.hnrrak
r€poned
fi.I di.r.ry pmrin
i .EiE sr5tm, Ar irfR@ cout.t bc d6M
dificmr lcvds ofdi.r.ry pet.in
Lhe ltndin8s
@nr.nt at which
of Srtvs ed
ndnm
for
oBinlD grbtllh.
Curte,o (taet,
pG6t culk @ d$ in sod.n.c wilh
varghcc (1936). Thcy
fin8..linss vilt!
30
qbn d
s
gbr1h of DlEor or!6 oklrrcd
abovc 45 percent though the eonomically oprinaldicbry plolein conrcnt was
Tne
Ih.y
I
p€rcnl
the tindinsr of Rautmdhy,
dE highgt SDwth in Cd,/a
.4rl,
ann Lab?o
and
nhitu
.dd l() !E@nr For.in dicc.
The smNrh p6rt€m
olfir
fth sFci6 rndd Fte
| .nd
I.l@ly
indicar.d $.1
.Iab.o tohira ad Cinhlns n'frS,/a n|d adjsrd io $6c .xl€nt ,gaiilr ncw cltrurc
nEdia (inleroiv. culrur.)
4 @mped
lo
calz .,/ta
md
nqlhm
h.ftnr of lhc dier wc
dcriv.d by th.s. rrc spei6. Thc fiDdings ofrh. lrudy suh€lniill. ihc
4!. (t993). Thcy Eponed
mdinh
nrisata qa! Labeo nhita on 30
BulB ofja!.d g
av.nsc inclEe in wiShl w.i.dainqt by
FcDt pmrin di.r cfiarnbrny g d.,
(197]) Eponed
conponlirly betcr Frfomsre ofcala.a,td in o intedle syscn rct
ihe supply of
.nifcirl
(z// ,l!m adificidl
fe.d.
Holew,
in
lh. p!!gr
Ct?r,,,r
itrg
mirJy on
srudy rh. b€Dtu c@i|r.n W Cana
te.d was vcry poor. Thc p6€nc. of Dbrot lie emyn6 i^
czlu (Rlmdc Md Kswalonmri.
I 9
higb prctein di.r lilic gmund-nur oit
76) sussesl physiotosiql c,pabitiry of
ro
u lD!
a
ote,
Th. orcrall f..d dd prot in.fljcicmic!
i. ph!*
rcsFcrivcry on lorv tFrein diet in
ce
Tle* dilrccd sisnilicorly fon
v"tu6 oforher
rhc
tsh
etla
I Enoincd
af Labeo tuhita
rh
bd
4 2 pe(.nt
fisl spei6. Ar
(0,25 .nd 0,64)
t.v€t of f.edin8.
4
Fr€
tevel oa
u9
fe'lin& led dd pbrcin
ddlined. Howv.r, lh.
emcicncics
ulue
nuimm 6hc of
vee @n/].n vrh Ldb@ r.hita
snd
or
rhc
f€.d and
tish
$dr
"p.ci6
trct in elfi.ienciq
ChrhiB wlgata, Esp..ti|.lv
on
all l@hents
(0-16 snd 0 17)
los ptolcin diet,
lh phase It the ov.rsll reed dnd piorcin cfuci.iciB sue ma*imum (0 29 and 0 59)
Bpdively
on high F otein di.ts
i\ .e
f..ding, Thee valree difieEd signifrcetly
.o//a. Al 4
FMt
ft.
f.d
(|
wss
6rl€
fon lh. vst!6 of
ed
poicin
r
lhc
Ldb.o rohita nid Catla
f..d dd prclcin ctrcienci€s w'E
tow pol.in di.t in
lik phs I Thw
e86,. I-hry Epon d oEl
35%
dd
delincd ftnd of f..d
FE.nt ldelorfdding
g.1.
in$ nlieald 5t 2 petcenl level of
lcvel of f..ding, th. bcsl valuc! of
Korded (0.r5 md 0.38) on high
eF€livclt,
Cn
ot
cs. ot Clnhin$ Nisola
.fiici*i.s
of l|.tFe fisn sFcies at 4
nndinr! coirci& with tlE Esults of Cailin
lowr f..di4
probin di.l h.d b.lcr f4d .mcicncv
dr
of*r
G%
vdc
bodv G'Chr r'
qtlish
(0.72) At hiSls fecdins
nLs
bodt eogfi0, fccd Gllci.ncv
vtls
of otltuh
Esulls Eponcd by Srcdpp. ard D.vaoj
(l9l)
contid dE litrdinss of pEsnt $udv
(4% of
M
Tn€y shovld
rhe hody
$!r Cdrl, .,rta
wieho
had poo@
fry fcd
.l 3ii fddirg
k*k
fc.l
4 8, 10,
BPidlv (0-50) Thc
12
snd 14
FEd
of
f.dd ud prcrciD .mci€ncies at hieh'r f'cdins lcrcl The re'd
dd pblein eflicid€ies \rw inpDv.d sigDilicdllv
hiSlEr
(4,
dali*d
md polein emcicnciq .r 2
P.'.e
sl$
ftd
!t low fddiic level nE
L.ding Bt. pdsiblv bc due ro
md'nm
utiliaion of sn.ll anounl of.vailablc di.l
Tbc reduclion in lccd ind prokin cflici.nci.s
obedsl rvi$ Zdr., rolta
Chthint Nisala u hit]ha t
nnd Cala
d
.nd
*iU
incrc$ing diclsry proteh
.otla EorEw this Ms
pol.h .mcicncis wF
of
'arc
lftlt poleit
Dol aue nr
@corded on
\"r
l20
dr€r
ir2
pcrccnt
f€sd .nd poteid
.m.i.nci6
dd
41. (1982)
(1t76)
Inc.
pcr.
lev.lot fecdirB whrtc ol4
s
w
0E
!@nr $!d, wE
Haqu. (1988)
ce
.ortrEdictory,
HokG,
im@ins lcv.l
lh.3. tEnd!
ql.1989). Th.y found rh.l
l.v.b of di.t !y
Ille
itrEiig rM&
But rlE
of Clrrluur DtaaL wit[
ed Tdcuchi g
diftdt
Cno
A d.
rcrb|cd thlr rad inr.I. .rd prct in .mciacy drio (r,ltr.)
*irh incEsins lq.l of di.t.ry ror.ir.
th. cfic.t of
d.crcril3 tr.nd ul
sinibr ro Lat.o rohha @n Atk uno.
d(Gasi wirl iicr@ing di.idrt pbr.in @rantl,
pol.in cfftci€ici.s in
tcvctoffccdinS rh.
nndinss
sqr
of fed and
of di.rary prctcin io
in lirc wirh rh€ Bulrs
of
f..d .flici@y of carF incees
my l.rls
pm|cin on thc fccd .nd
ro rh. conclusion rhd
p6t
in
cflici.'Ei6 v.ry
slei6,
lcordina b ln. fish
Thc b.st prclcin
dd.4rry @nEiio!
cllici.tuics !ft16
Phe
I
*E
@l.ular.d
c
l*l of f..din8 *hiL
4 p.E.nt lal of fcdins, Crrfirt 6 #'l4td ito*Ed tlE hiSlEsl rlF of 5.4 md 7-5
'|
8.1rnd
10.9 on low
pret.in dict
on low and high pdLin dieb
olcdll
bcst polcin
poinl of
ob$.lion
thN
fish
sFri6
ed
ir
ftrF
in(g
6Ftiv.ly,
finhr nFnalhd.d
val$s q.E
9.6 and | 4.6 for
rl
level of fcding,
Frc.nl
.rmiring
by
wlEE .Lposilion oi pbtcin
F.
r
Horevd, rtalbtic.l @lysb shorql El
@nrcRion .mci@ci.r in
As Esdds tlE Plorcin .J|d
2
ol Lab.o ahXa.r2
ed
tal
ca
thc
C,rr,t@ Df4,/a nG
d.|ji on hody @dposnion
of
@ dr. w@ coep.6liv.ly hish.r
.ners .onERion .mci.nciqs u
ter
plE. il,
rh.
bcsr
Cial'Drr rr,8al, on low .rd ligl prlcin di.t rsp(tiv.ly
Wh.@ .t
of prctcnr and .n.rsy w.E 6.4
sd
8.1
4 p.Mnr lcvrl of fdding, th. muinum vrlucs
.lso
tu
c4
ot
C/plt 6 rrrSala oi loo pe|.ir
'
di.l. Th.daliuingtEnd
otprcLin.
B s.
Ph6e L
te.diq
Bllts
The
oblained
s undr
ftrc
pr*ni
ed
dE
fceding
nlo of
5% of wel body \vcisht.
study, higlEr values of p@r.in and crergy conv€rsion,etlicidnc,cs
w.E low.i Sinilar aeid
hidrd sd
..
lo* prukin did wh€qs oi m€diun &d hid pol.in
ob$ryed on
Th€y
porcin 5nd .neret convcrsion .mci.rci.s at lhc lowcr feedhg dlc
(2% or1rct body q.igho dnd minnnnm ellues
In the
snvdion cftci.rcie! tl4 pcrc.rl tcvctof
rgn€ndl with rlE fiodiiss or cattih q 4. (t986)
w€rc in
nrxinln
ener$,
ltt
@
rhoM by R'jbsBhi e!a!.
lo@i prcroi cfnc'.fty 610
50,4 p€rced crude prot€in.
cu.d
(1989). Th€y Gpon.d
in ohu
qp+tivelt. Howwr,
thc
dieis the v6lues
w$ ;i.k conrdinS
r*llls \we
dd
19 18
@nlraddory lo
$e findrn$ ofFag.rlud g 4!. (198J). fhcr conp{E.l lh. eh(t ol15% (LP, dnd 519.
(HP) prct in di.t on FoGin and .a.r&| 6nrc6ion .flrcj.nciF in
dd EFrl.d
rI.
lhal
hign porcan l.vcl.
ulB
of troLin anl.n€tgy
Mmy 4
gl. (197?)
Juqil.
Coho
@r@ion.Ilicidis sw
de .gr..d
sdno.
hiShd al
wnn dE FporL of F.g.rlund
q d.
(lgEl) in which th.) sl$rcd ihpov€d potein @nvdion qho dictirv polei' 9A
iocEsi tom
25% lo 15%
fingcrlinS carfish.
di0cEnl polch
dd tuilq
Th* ob*rydiN
EquiEm.
for
$ggca
supFrtcd lhe
0r
nighq Prelein EqliEm.fl for
vi.e th.l
diffdt fth spais
hdlt
bd @nwsion.fiicimy
BODY COMPOSITION
Like
afaecred by
dt
body
wi8lt
of fish, lhe body conporilion of thEc fish species w.E atso
thc lcwls of feeding. At highs fe.ding
de
in
Phe I, noisllt
contcn*
d6da3.d *hile Prollii, f.l md sh conlcnti
Uflld Phe
ohly lsh contenG decGss.d.
cont..B d.ccAed
enit
Ar
ed sh
lowt
acerting
@nt ntt
d4Esd
*hil. potcin ed
conrcnb
d.,
nr@*d
code ! deqe*d while
tish a12
fitrdi4s
wE
d
I
ol Lzhto
tlhib
enb inc@s€d wniL prcl'in' fat
se
p€niallv in
hieher f'c<lins
of
enrt increed
l'd"' ro'ia
'gtEoar
id''
o
whee Prol.in
wi! fi' suh
of Gtrlin
g
bodv noi3tuE' porein a|ld dt)
Thc low fal
dd polein nccrnuhnon 'n
nad lvhich Eldded
mov b' dw b f@d $pplv b'low its
lcv'l
fcdine
Frc6t
pbgesire ftducdon in fat
lnd Vsnstone, 966) A@rdins to
dd sktu lo nobili4
netrly lsed up After diat,
lh€
shin asuled
s
L'v'
(1980)' fie fsh
pbl€in
is
lipids fron
tis
tndsl
$urce of
fie
has De€n
phce Such a
ilizcd' water mov€s in lo mr' ns
Th' ftnds of noi$ure @nlcds
iFEs$d wacr @nrcnts of th' bodv
'n
'l-he ovedll higher
th€ tbove
vo*ds
pceDltsc of prclein md flr mong thEe
frmd in ce of Cr?r,ti,$ tr'Sdlt @
hiSh prclein
difereuoes anong sFcics
w'e signifimt
dulc aport'd
bv
llh
did al bo$ fc'dhs
md lrigh pol.in dieB
$e dific!.nes mong low, m€diu
in |nE Nitlr
and prctein rcserves (Pdker
tl 66r coEm'd
Protins onlv *htn
n$l'
offish body s.r€ suppon.d bv
N
moisluie cnd asn
@
misn'€ dd 4{r
dc'pi it
onl€ s inc@trl
fai
th. StoMl of lith md dus'd a
liyer
uldd phd
\t|@
(1986). Th.y epolted lhdt
ol Catla caltt \ube€
higt'r f'€ding rale
Phe l, moistw co
un&r
fat conl.Dls d'cE@d
T[*
cu
'ar
as
dt
ll
in
ontcnls iNrc&rcd qc'pt in
l,Frein Dd
wh.E pot incont ol5d.ct
incred qepr
wt'
w*
I'vtls Hos?v'r'
non_sisni[csl wbile lhe
rbr th€ bodv @rPosition Th'
hltnm ud
specics
Pa*nr nrdines
cffecl ot
She(t (19s0) sho conclded rh!|
tza
difi.r.nr t@uing lrd fading rEgic on body @dposirion ofcdla catta Labeo rchita
ed Cttinut .arpiow mr sigrifimtl, di|I.Ml. Hov*r,
lEinenB dtoncd
by Javed (19E3)
sc
not in
ehcda sF.i.s diff@nc6 obsrcd by fiin
fircu
e
rh. djfrcEnc., of
of tuxlinF ofihc tE$nr study
.8Eddr
with th. prcsnl srudy.
Zenk. g sl.(1984) olso Epon€d $dl dicrlry cruds prot.in cod€nrs over 4t.3%,
nei$d rh. For.in qonrc smr far conrdrr ofrh.
rh.a qa !
fat
d.c|!e
conlat! of thc
of rhe prer.in
c.|!s
@
mr3 ldd
cd6
sierifi@tty but b.low 41.3%
dhdr.Fudy v.ry hid ircce
of bodt
@Fposition
of lhE tish speics
sisnifionrly negativ. conel.rion among mobrula, peLin
at 2 pcrc. nt
le
in $e
ofcarp.
Corcl.tion Mlrsis
rorrd
chanSe
dd
far
rcv@l.d
@nr.$ ot ldbeo
rcl of f.qljne exccpl lh. cor€l rtion b.rw.n noistuc dnd ash lvhich
Ms nor- riSnilic&t. At 4 p€Mt l.wl of &cdin3, th. .orelation @ftici.nr3 ahong
nroistur,
pot.ii, f.t &d sh v@ sisDifi@rly rEgdirc qc.pr
$,bich shoe.d polnivc
oftlalion.
far rnd .slr conr.nb of
@mci€
s mons hoisluE, pror.in,
Catlu catlz
dnwri loDq{har sihild
Thc cofrlatioi
Corlrrr wiAala s4
p^r.tn of Lubeo rchita, \he pesent work subsidlirt
d
lh.
nndinCs of
rhE fisfr sFci6 bb?o rohita, Cdlu.dlo
Mumy el d.
(|
977) tlE sigtrifc&t ftearivc
moisturc conr.nr! dcnoosrErcd ih. diluting
Tle corclation .oem€i€nls
atd
rartd
1988), wbo
ed rol.l fia
dl
Cinh,t6 D/,Sdd _ Acoding
ro
.cl.ion tawd c!M$
.ll.ci
(
for
Eponed signili@dy iegative orclation ohorg w.tci, rot l pnrciro
rhc
nd noisl@
pror.in
of@a
lipidr
on
litid lrd
w.rcr
berween prctcin and tat coftents
cd6s
hds,
ol Lubaa rchito
Fn.nr.d sigilictrtrtly posirive but fot Cirthi'M ndgala .nd Catla.a?id rh. corclalioi
tx
Nas negafttly
rcFsigdficinl al
l)crce lewl ol rbcdin! Al 4 Pc,ce.r lcvcl of t_edi'&
2
contlaion b.rweeo prct in and fat coot.nts of all
posnn€ and
$b
$re
t clo* inle.nltlio$hip dong
rcvcat.d
findiner sutpon dE rcsulls of Z.nkr q d. (1984)
dd
EE
3ignificanllv
ndri'nli
The Prcsent
tun spai6
ncal
JaEd (1988) Thcv ob*dcd rlch
@ftlation b.hrEtr Lr and pmch in CWti,B c@Pio Lab'o tohitd. Cltla 'dla
Th. @frlatior *flicient!
ht Ctuhin8
tuigala
b€t*d 6h ed
sA Cdld atla
wt$d
porein
wF sisnifiddv "gatire
for l'tbeo rot'nd @ftlatlon
w
noF
pe@nt lev'l of L'ding s6
tignificdl !t 2 Fcnt ledina leEl This @ltrion 'r 4
u\d 98itu' fq Citthitd Dtlrata howevd lor
signif!61 but mslnv. for 12, @
'ohita
Cdlo .a,ld @ftldion
Th. corcladot
6
n saliwlv mn'sigtrin@t
b.t$i
pGnisly non{i8rifi@r
for
L
'!h
beo
and fat al 2
rohita Nd CaId
it w6 posilvely sigri6tu! shil'
posirively siSdlicsnt aor
lgnin@r
b€en tlu. lo
Such
all $rc'
diff.Ens
pd'61 l4cl of f"ding
lh€
corehio!
fth s,e'i*
in bodv conposilion
'atla
ar 4
'xc'Pt
B
lound to be
*rE@ fil Cirhisd
ndgata
F@nt lev'l or f'cding
Cal'
son8 th@
'otla
w
which hsd tro$
nsh sD'cies nigbl hale
inknsiv' svst€r' Mor'over'
diI1irflr palkm of 8ro$4h Frfornue udo
otlrr @!$s of ldiations in
bodv conpNilion
colld be lhe f?id iniak'' nct'bolisnl
habits oflish'
cflicicocy, ereqv e{pendirlt ud fFdins
A!
ll
re
nsh
coclarion .nalvsis of bodv comPosition of $@
vas conc€nE<l, fcw vari.rion!
*te
Ecotded This
v.gerflble prokin sourccs frighl be sliShtlv
in
u€nced
t'vak
species
undd
phe
thri dtio of animals 10
$e bodv conpositun ot
tfiN
fish
sF.i.r. lltc difl.Ena
oa
@
corcl.lion bd@n hoi.toE {nd pFlcin
of
found nr c6c
Cdla.a,/d which chrB.d fom podiv.ly non*isnifi@l to rceBlircly sistificst dt 2
percent
l.vcl of fe€ding. Sidilar
conGnk
otcal&
cr?/z- As lh€
change was alm
and
I97D. Tn€ Edd of
w6s
lik. phAe
@clalion
Labeo
m
poLin @nrcn|s
shoeed
I
ch48td liom
sigtili@i. At 4 F@di level of
w
The corclalion
non{igniti@t
ofrt{rion baq@ motlw ..d sh
vhile L4beo tohita
8d
f€eding,
bctren
idca of
NoislLre
Muffit c! al
@nt'n|s of Cirtuinut ntiAalu
Cbl/4 cala show'd sonc veialioh The
ch&gql fom reSaiivclv significdl !o positirclv non'ligrifcMl
tuhib $nqr6}ilrb Cai4 ddrl! tlE c@ladon
ol .oftldlion cocmci.n$ belsecn polein
$€
lollow.t lhc s@ p'll'm ot ph'*
wu similar to phe l. This tudhd @nfidcd 0E
aod tar contots
(
'fti! @ftl.tion *a
show.d veidion.
n.gativc non+ignificui lo Positively
I exced Z?6eo,orirz which
far
@clation @flici.nts b.l@n noisturc dd 6h co.ten|l
w.e .orcdn//.. ttb.o rohirt
coftl.tion bcncr moislut
obsd€d bawsn ,noisnrrc and
sd fd
b{ee
i' ca'
changa
'ew
level of feding
3igniliclnl' A
@nt'nts ot 2
pe&e
\E€ .l$ Kotd.d in ce of Ctthi,us ndgold ed Cald 'atla \,'iilc Lab'o tohiti
fll cod€nls
Gv.aled lh. s.lr€nd otphN I Th. €orclslion co'fficienl ofpol.in '!d
ol Citthtn8 nngola
*d
Catla./,i4 reG chmsed fon n'gativelv non_sig'iUca to
posiiiv.ly siSnilics This posilivclv
sieni{ie trdd
clos inlcFFlatonship mo.s nqt tur''nr! At 4
,
thc slEnglh to
SavG
p'G
tcacl of f€cdins' thc lEnd
corelarion ccmcic.b betsedn ptoGin md l6t conl? s of
fotlo\rcd
phe I d.€!r cal/ ../la
which
fie id" ot
thF
cho8'd ton sisnificdt
'
ot
fish spccid 1wE also
1o
rcn_!'81:iti@tr'
116
TIE
of
ins
cmLri@
6.i*d
cqr.lotion
of
rta
A1 4
S*
in
st
onrcnl5
oflar.o
toitird .r 2
t
c,rri,B
@.1
mis,/d
b
to p.6iIivc,
t.v.l of f.atinr; $. c@l.rion b.t$@ P'd.in od sh
w
sinilr s ph4l wlil.
of
,r&o.,rr. !E |tad E
mD-5isninc&t .nd of Cd,l4 calz noD
lslly. thc lted of cofrl.!@
sD.ci.s
od
sc s iafi,lt I v\ih inChUM 6.1gda atd Catl, cdtl, thc
t@l
shjfi.d fioD sianitidn
t
not
w.bDgd fFn 3isnifiol lo ootrrigift€at.td t@ rg.liE
Gp.ctiElt.
srdlr
ba*6
@dnd ! bct*er frt ed 6h
sl$rtd cohpdttivcly ooE widnN d D.(l
IGY€lt
mn'!i8ni66l
conllnts
of&.diD€;
to